Paleoanthropology: The Study of Human Lineage
[1] The lineage of which we have to speak in the present chapter embraces all fossil and genetic evidence found anywhere, or at any time, beneath the earth’s surface or within its populations. Particular milestones in this deep history have, of course, their specialized nomenclature. Australopithecus afarensis, the poster-fossil for early bipedalism, is a form of hominin that has played a very[2] influential part in scientific narratives; hence the separate mention of “ape-men” in the heading of this chapter.
While there are many selective pressures that help to shape the arc of evolution, the most potent of them all is environmental change. Let us see what happens when a shifting climate alters the landscape of a tropical forest, for example. If the ecosystem transformed in a smooth, linear fashion over such a terrain, its pressure might nudge a species along a predictable path, but would not fracture a population into distinct branches. Such gradual adaptation, due to the consistent component of environmental stress, does, of course, occur, and its effects are strikingly visible in the incremental dental or postural changes observed in the fossil record. [3] These anatomical shifts clearly illustrate how steady pressure refines a species over eons. All environments in the ancient world were, however, full of volatility and catastrophe, and in many epochs, as during the rapid oscillations of the Pleistocene ice ages, there were strong selective shocks, sometimes extending to mass extinctions.
All species are bound by their genetic heritage,[4] and, contrary to a teleological view, never truly “strive” toward a predetermined form. Speciation may occur in isolated refugia, through the fragmentation of populations by rising seas, or it may be sparked by novel behavioral innovations, but a new hominin lineage emerges only because the ecological niche is changing with it; and, in a stable world, all species remain more or less static in their form. The rate of its evolutionary change depends upon the generation length, and upon the size and stability of the breeding population. [5] Other things being equal, the smallest, most pressured populations change most rapidly. Exceedingly large and widespread groups, even without intense selection to drive it, require hundreds of thousands of years to show significant morphological divergence from their ancestors.
Upward trends in cognitive complexity suffice to distinguish the genus Homo[6] uniquely every few million years, and migratory movements carry the same bands far and wide over the earth. The dispersal of populations by foot and raft leads to some results of remarkable interest, genetic as well as cultural. [7] In the first place, far-reaching differentiation in morphology and technology are brought about by this process.
[8] 1. Thus, in Indonesia, a distinct and diminutive hominin, Homo floresiensis, persisted in isolation long after its presumed ancestors had vanished elsewhere. 2. It is believed to have become adapted to the limited resources of its island environment. 3. Less extreme examples of this adaptive radiation are found on many other landmasses, including the whole of Eurasia for the Neanderthals. 4. Another effect of geographic dispersal is the hybridization of lineages. There is now clear evidence of genetic interchange between different hominin groups, so that the genome [9]of a particular ancient individual, [10]as a result, is not purely one species but a mosaic of several. Lastly, the presence of multiple, coeval hominin species in the atmosphere of competition, whether derived from a common ancestor or otherwise, had various profound and decisive effects upon the cognitive and social capacities we associate with modernity and modifies, in numerous ways, the conditions of human emergence upon [11]the planet.